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Organism
  Animalia
      Arthropoda
            Insecta
                    Coleoptera
                            Scarabaeidae
                                     Anomiopus
                                     Aphodius
                                     Ataenius
                                     Ateuchus
                                     Bdelyrus
                                     Canthidium
                                     Canthon
                                     Ceraspis
                                     Chlorota
                                     Copris
                                     Cryptocanthon
                                     Cyclocephala
                                     Deltochilum
                                     Dichotomius
                                     Eurysternus
                                             Eurysternus caribaeus
                                     Golofa
                                     Hemiphileurus
                                     Heterogomphus
                                     Isonychus
                                     Lagochile
                                     Macraspis
                                     Macrodactylus
                                     Megasoma
                                     Ontherus
                                     Onthophagus
                                     Pelidnota
                                     Phyllophaga
                                     Platycoelia
                                     Plectris
                                     Pucaya
                                             Pucaya castanea
                                     Stenocrates
                                     Sylvicanthon
                                     Uroxys